Coast Douglas-fir
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| Coast Douglas-fir Conservation status: Secure | ||||||||||||||
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| Missing image Pseudotsuga_menziesii.jpg Coast Douglas-fir foliage | ||||||||||||||
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The Coast Douglas-fir (Pseudotsuga menziesii subsp. menziesii) is an evergreen conifer native to the coastal regions of western North America, from west-central British Columbia, Canada southward to central California, United States. In Oregon and Washington its range is continuous from the Cascades crest west to the Pacific Ocean. In California, it is found in the Klamath and Coast Ranges as far south as the Santa Cruz Mountains, and in the Sierra Nevada as far south as the Yosemite region. It occurs from near sea level along the coast to 1,800 m in the Sierra Nevada. Further inland, Coast Douglas-fir is replaced by the related Rocky Mountain Douglas-fir (Pseudotsuga menziesii subsp. glauca).
Coast Douglas-fir is a very large tree, the second-tallest in the world (after Coast Redwood). Trees 60-75 m (200-250 feet) or more in height and 1.5-2 m (5-6 feet) in diameter are common in old growth stands. The tallest specimen is the "Brummit Fir", 100.3 m tall, at East Fork Brummit Creek in Coos County, Oregon, the stoutest is the "Queets Fir", 4.85 m diameter, in the Queets River valley, Olympic National Park, Washington. It commonly lives more than 500 years and occasionally more than 1,000 years.
The bark on young trees is thin, smooth, gray, and contains numerous resin blisters. On mature trees, it is 10-30 cm thick (4-12 inches) and corky. The shoots are brown to olive-green, turning gray-brown with age, smooth, though not as smooth as fir shoots, and finely pubescent with short dark hairs. The buds are a very distinctive narrow conic shape, 4-8 mm long, with red-brown bud scales. The leaves are spirally arranged but slightly twisted at the base to lie in flattish either side of the shoot, needle-like, 2-3.5 cm long, green above with no stomata, and with two whitish stomatal bands below. Unlike the Rocky Mountain Douglas-fir, Coast Douglas-fir foliage has a noticeable sweet fruity-resinous scent, particularly if crushed.
Pseudotsuga_menziesii_cone.jpg
The mature female seed cones are pendent, 5-11 cm (2-4 inches) long, 2-3 cm broad when closed, opening to 4 cm broad. They are produced in spring, green at first, maturing orange-brown in the autumn 6-7 months later. The seeds are 5-6 mm long and 3-4 mm broad, with a 12-15 mm wing. The male (pollen) cones are 2-3 cm long, dispersing yellow pollen in spring.
In forest conditions, old individuals typically have a narrow, cylindric crown beginning 20-40 m (65-130 feet) above a branch-free trunk. Self-pruning is generally slow and trees retain their lower limbs for a long period. Young, open-grown trees typically have branches down to near ground level. It often takes 70-80 years for the trunk to be clear to a height of 5 m (17 feet) and 100 years to be clear to a height of 10 m (33 feet). Appreciable seed production begins at 20-30 years in open-grown Coast Douglas-fir. Seed production is irregular; over a 5-7 year period, stands usually produce one heavy crop, a few light or medium crops, and one crop failure. Even during heavy seed crop years, only about 25 percent of trees in closed stands produce an appreciable number of cones. Each cone contains around 25 to 50 seeds. Seed size varies; average number of cleaned seeds varies from 70-88/g (32,000-40,000 per pound). Seeds from the northern portion of Coast Douglas-fir's range tend to be larger than seed from the south.
Ecology
Coast Douglas-fir is the dominant tree in the Pacific Northwest, occurring in nearly all forest types, competes well on most parent materials, aspects, and slopes. Adapted to a moist, mild climate, it grows larger and faster than Rocky Mountain Douglas-fir. Associated trees include Sitka Spruce, Sugar Pine, Western White Pine, Ponderosa Pine, Grand Fir, Coast Redwood, Western Redcedar, California Incense-cedar, Lawson's Cypress, Tanoak, Bigleaf Maple and several others. Pure stands are also common, particularly north of the Umpqua River in Oregon. Shrub associates in the central and northern part of Coast Douglas-fir's range include Vine Maple (Acer circinatum), Salal (Gaultheria shallon), Pacific Rhododendron (Rhododendron macrophyllum), Oregon-grape (Mahonia aquifolium), Red Huckleberry (Vaccinium parvifolium), and Salmonberry (Rubus spectabilis). In the drier, southern portion of its range shrub associates include California Hazel (Corylus cornuta var. californica), Oceanspray (Holodiscus discolor), Creeping Snowberry (Symphoricarpos mollis), Western Poison-oak (Toxicodendron diversilobum), Ceanothus (Ceanothus spp.), and Manzanita (Arctospaphylos spp.). In wet coastal forests, nearly every surface of old-growth Coast Douglas-fir is covered by epiphytic mosses and lichens. Its rooting habit is not particularly deep, the roots tending to be shallower than those of same aged Ponderosa Pine, Sugar Pine, or California Incense-cedar, though deeper than Sitka Spruce. Some roots are commonly found in organic soil layers or near the mineral soil surface.
Coast Douglas-fir seedlings are not a preferred browse of Black-tailed Deer and Wapiti, but can be an important food source for these animals during the winter when other preferred forages are lacking. Douglas-fir seeds are an extremely important food for small mammals. Mice, voles, shrews, and chipmunks consumed an estimated 65 percent of a Douglas-fir seed crop following dispersal in western Oregon. The seeds are also important in the diets of the Pine Siskin, Song Sparrow, Golden-crowned Sparrow, White-crowned Sparrow, Red Crossbill, Dark-eyed Junco, and Purple Finch.
The Douglas squirrel harvests and caches great quantities of Douglas-fir cones for later use. They also eat mature pollen cones, developing inner bark, terminal shoots, and tender young needles.
Mature or "old-growth" Coast Douglas-fir is the primary habitat of the Red tree vole and the Spotted Owl. Home range requirements for breeding pairs of spotted owls are at least 400 ha (4 km² / 1,000 acres) of old-growth. Red tree voles may also be found in immature forests if Douglas-fir is a significant component. This animal nests almost exclusively in the foliage of Douglas-fir trees. Nests are located 2-50 m (6-160 feet) above the ground. The red vole's diet consists chiefly of Coast Douglas-fir needles.
In many areas Coast Douglas-fir needles are a staple in the spring diet of Blue Grouse. In the winter, porcupines primarily eat the inner bark of young conifers, especially Douglas-fir. Douglas-fir snags are abundant in forests older than 100-150 years and provide cavity-nesting habitat for numerous forest birds.
The leaves are also used by the adelgid Adelges cooleyi; this 0.5 mm long sap-sucking insect is conspicuous on the undersides of the leaves by the small white "fluff spots" of protective wax that it produces. It is often present in large numbers, and can cause the foliage to turn yellowish from the damage in causes. Exceptionally, trees may be partially defoliated by it, but the damage is rarely this severe.
Uses
Coast Douglas-fir is one of the worlds best timber producers and yields more timber than any other tree in North America. The wood is used for dimensional lumber, timbers, pilings, and plywood. Creosote-soaked pilings and decking are used in marine structures. The wood is also made into railroad ties, mine timbers, house logs, posts and poles, flooring, veneer, pulp, and furniture. Coast Douglas-fir is used extensively in landscaping. It is planted as a specimen tree or in mass screenings. It is also a popular Christmas tree.
External links
- Arboretum de Villardebelle - cone photos (http://www.pinetum.org/cones/PScones.htm)
- Gymnosperm Database: Pseudotsuga menziesii subsp. menziesii (http://www.botanik.uni-bonn.de/conifers/pi/ps/menziesii2.htm)
- [1] (http://www.fs.fed.us/database/feis/plants/tree/psemenm/index.html)de:Douglasie